Sandy Kawano

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Sandy Kawano

Assistant Professor of Biology


Email: Sandy Kawano
Office Phone: 202-994-0017
SEH 6880


Comparative biomechanics and physiology; functional morphology; water-to-land transition in vertebrate evolution; natural selection; musculoskeletal function.

Please refer to my website for recent research information.

1.     Kawano SM. “Selection Gradients.” In Oxford Bibliographies in Evolutionary Biology. Ed. Douglas Futuyma. New York: Oxford University Press, forthcoming.

2.     Moody KN*, Kawano SM*, Bridges WC, Blob RW, Schoenfuss HL, Ptacek MB. 2017Contrasting post-settlement selection results in many-to-one mapping of high performance phenotypes in the Hawaiian waterfall-climbing goby Sicyopterus stimpsoniEvol Ecol. 31(4): 489-516. * = co-first authors. 

3.     McInroe B*, Astley HC*, Gong C, Kawano SM, Schiebel P, Rieser JM, Choset H, Blob RW, Goldman DI. 2016. Tail use improves performance on soft substrates in models of early vertebrate land locomotors. Science 353(6295): 154-158. doi: 10.1126/science.aaf0984. * = co-first authors. 

4.     Kawano SM, Economy DR, Kennedy MS, Dean D, Blob RW. 2016. Comparative limb bone loading in the humerus and femur of the tiger salamander Ambystoma tigrinum: testing the “mixed-chain” hypothesis for skeletal safety factors. J Exp Biol 219: 341-353. doi: 10.1242/jeb.125799. Data:

5.     Kawano SM, Blob RW. 2013. Propulsive forces of mudskipper fins and salamander limbs during terrestrial locomotion: implications for the invasion of land. Integr Comp Biol 53(2): 283-294. doi: 10.1093/icb/ict051. 

6.     Bohórquez-Herrera J, Kawano SM, Domenici P. 2013. Foraging behavior delays mechanically-stimulated escape responses in fish. Integr Comp Biol 53(5): 780-786. doi: 10.1093/icb/ict031.

7.     Kawano SM, Bridges WC, Schoenfuss HL, Maie T, Blob RW. 2013. Differences in locomotor behavior correspond to different patterns of morphological selection in two species of waterfall-climbing gobiid fishes. Evol Ecol 27: 949-969. doi: 10.1007/s10682-012-9621-z. Data:

8.     Schoenfuss HL, Maie T, Kawano SM, Blob RW. 2011. Performance across extreme environments: comparing waterfall climbing among amphidromous gobioid fishes from Caribbean and Pacific Islands. Cybium 35(4): 361-369.

9.     Blob RW, Kawano SM, Moody KN, Bridges WC, Maie T, Ptacek MB, Julius ML, Schoenfuss HL. 2010. Morphological selection and tradeoffs between predator escape and climbing in Hawaiian gobies. Integr Comp Biol 50(6): 1185-99. doi: 10.1093/icb/icq070.

Ph.D. Biological Sciences. Clemson University, U.S.A. 2014.

B.S. Evolution, Ecology, and Biodiversity. University of California, Davis. 2008.

A.S. Biological Sciences. De Anza College. 2005.